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The Lower Silurian period-The Silurian rocks generally-Limits of Lower and Upper Silurian-General succession, subdivisions, and characters of the Lower Silurian rocks of Wales-General succession, subdivisions, and characters of the Lower Silurian rocks of the North American continent-Life of the period - Fucoids - Protozoa - Graptolites - Structure of Graptolites - Corals - General structure of Corals - Crinoids - Cystideans - General characters of Cystideans - Annelides - Crustaceans - Polyzoa - Brachiopods - Bivalve and Univalve Molluscs-Chambered Cephalopods-General characters of the Cephalopoda-Conodonts.

Whilst all the sub-kingdoms of the Invertebrate animals are represented in the Lower Silurian rocks, no traces of Vertebrate animals have ever been discovered in these ancient deposits, unless the so-called "Conodonts" found by Pander in vast numbers in strata of this age [15] in Russia should prove to be really of this nature.

[Footnote 15: According to Pander, the "Conodonts" are found not only in the Lower Silurian beds, but also in the "Ungulite Grit" (Upper Cambrian), as well as in the Devonian and Carboniferous deposits of Russia.

Should the Conodonts prove to be truly the remains of fishes, we should thus have to transfer the first appearance of vertebrates to, at any rate, as early a period as the Upper Cambrian.

On the contrary, whether the so-called "Conodonts" should prove to be the teeth of fishes or not, we are justified in expecting that unequivocal remains of this group of animals will still be found in the Lower Silurian.

These are numerous, but, with the exception of the still dubious "Conodonts," belong wholly to the groups of the Ganoids and the Placoids (including under the former head remains which perhaps are truly referable to the group of the Dipnoi or Mud-fishes).

The Fishes are the lowest class of Vertebrates, and they are the first to appear, their first certain occurrence being in the Upper Silurian, whilst, even if the Lower Silurian and Upper Cambrian "Conodonts" were shown to be the teeth of Fishes, there would still remain the enormously long periods of the Laurentian and Lower Cambrian, during which there were Invertebrates, but no Vertebrates.

Conodonts, 114, 131.

Some of these belong to Cyrtoceras, which only differs from Orthoceras in the bow-shaped form of the shell; others belong to Phragmoceras, Lituites, &c.; and, lastly; we have true Nautili, with their spiral shells, closely resembling the existing Pearly Nautilus. [Footnote 14: This illustration is taken from a rough sketch made by the author many years ago, but he is unable to say from what original source it was copied.] Whilst all the sub-kingdoms of the Invertebrate animals are represented in the Lower Silurian rocks, no traces of Vertebrate animals have ever been discovered in these ancient deposits, unless the so-called "Conodonts" found by Pander in vast numbers in strata of this age [15] in Russia should prove to be really of this nature.

The whole of this difficult question has thus been reopened, and we may yet have to record the first advent of Vertebrate animals in the Lower Silurian. [Footnote 15: According to Pander, the "Conodonts" are found not only in the Lower Silurian beds, but also in the "Ungulite Grit" (Upper Cambrian), as well as in the Devonian and Carboniferous deposits of Russia.

Should the Conodonts prove to be truly the remains of fishes, we should thus have to transfer the first appearance of vertebrates to, at any rate, as early a period as the Upper Cambrian.] Page 115 CHAPTER X.

On the contrary, whether the so-called "Conodonts" should prove to be the teeth of fishes or not, we are justified in expecting that unequivocal remains of this group of animals will still be found in the Lower Silurian.

These are numerous; but, with the exception of the still dubious "Conodonts," belong wholly to the groups of the Ganoids and the Placoids (including under the former head remains which perhaps are truly referable to the group of the Dipnoi or Mud-fishes).

The Fishes are the lowest class of Vertebrates, and they are the first to appear, their first certain occurrence being in the Upper Silurian; whilst, even if the Lower Silurian and Upper Cambrian "Conodonts" were shown to be the teeth of Fishes, there would still remain the enormously long periods of the Laurentian and Lower Cambrian, during which there were Invertebrates, but no Vertebrates.


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